In this episode we gently debate subspecies, and then dive into several snake papers, from their anomalously extensive and rapid diversification, to a recent taxonomic change. Regrettably the audio on this episode was affected by a technical error, so it is far less than our usual quality (Ethan and Hiral are a little quiet, sorry!). Hopefully it doesn’t affect the listening experience too badly!
Remember that you can watch the full episode with video at youtube.com/@squamatespod!
Episode notes sometimes get clipped on your device or by your podcast provider; for full notes and references, go to squamatespod.com
Check out Ethan’s new book, The Unfamiliars!

Episode Citations
Vences, M., Köhler, J., Scherz, M.D., Hutter, C.R., Rabe Maheritafika, H.M., Rafanoharana, J.M., Raherinjatovo, H., Rakotoarison, A., Andreone, F., Raselimanana, A.P. & Glaw, F. (2024) Four new species of forest-dwelling mantellid frogs from Madagascar allied to Gephyromantis moseri (Amphibia, Anura). Spixiana, 46(2):297–319.
Scherz, M.D., Rudolph, J., Rakotondratsima, M., Ratsoavina, F.M., Crottini, A., Andreone, F., Glaw, F. & Vences, M. (2024) Molecular systematics of the subgenus Gephyromantis (Phylacomantis) with description of a new subspecies. Zootaxa, 5446(2):205–220. DOI: 10.11646/ZOOTAXA.5446.2.3
de Queiroz, K. (2020) An updated concept of subspecies resolves a dispute about the taxonomy of incompletely separated lineages. Herpetological Review, 51(3):459–461. [Mark incorrectly gave publication date as being 2021]
de Queiroz, K. (2007) Species concepts and species delimitation. Systematic Biology, 56(6):879–886. DOI: 10.1080/10635150701701083
Title, P.O., Singhal, S., Grundler, M.C., Costa, G.C., Pyron, R.A., Colston, T.J., Grundler, M.R., Prates, I., Stepanova, N., Jones, M.E.H., Cavalcanti, L.B.Q., Colli, G.R., Di-Poï, N., Donnellan, S.C., Moritz, C., Mesquita, D.O., Pianka, E.R., Smith, S.A., Vitt, L.J. & Rabosky, D.L. (2024) The macroevolutionary singularity of snakes. Science, 383(6685):918–923. DOI: 10.1126/science.adh2449
Strong, C., Scherz, M.D. & Caldwell, M.W. (2021) Deconstructing the Gestalt: New concepts and tests of homology, as exemplified by a re-conceptualization of “microstomy” in squamates. Anatomical Record, 304:2303–2351. DOI: 10.1002/ar.24630
Losos, J. B. (2017) Improbable Destinies: Fate, Chance, and the Future of Evolution. Riverhead Books, New York City NY, USA. 384 pp.
Datta, D. & Bajpai, S. (2024) Largest known madtsoiid snake from warm Eocene period of India suggests intercontinental Gondwana dispersal. Scientific Reports, 14(1):8054. DOI: 10.1038/s41598-024-58377-0
Zaher, H., Trusz, C., Koch, C., Entiauspe-Neto, O.M., Battilana, J. & Grazziotin, F.G. (2024) Molecular phylogeny and biogeography of the dwarf boas of the family Tropidophiidae (Serpentes: Alethinophidia). Systematics and Biodiversity, 22(1):2319289. DOI: 10.1080/14772000.2024.2319289
Vences, M., Guayasamin, J.M., Miralles, A. & de la Riva,