Willow Pipp, Simon Maker, Fennel Wispywhisks, and Squeaky McSquirmels face off against the wicked Princess Maybelline, attempting to secure the Staff of Suetonius from its crystal cocoon. Afterwards, once again mounted on Sir Snuffles, they journey to the Northern Marches, where Lord Balthazar and his Pierats dwell in the city of Chimneyhaus. An epic battle ensues, and the lowly mice of Bernpyle must rise to the occasion and prove they truly belong among the renowned ranks of the Mausritter.